Location: Home > Scientist > Faculty and Staff
Faculty and Staff
Academic title:
Postal Code:
Subject categories:
Mailing Address:
88 Xuefu Road, Kunming, 650223, Yunnan, China

Nationality:          Chinese

Date of Birth:        8,Octomber.1962


Ph.D   Yunnan Universityand Frankfurt University of Germany(1998-2001).

Bachelor  Yunnan University (1979-1983).


 Chemical ecology and Honeybee science




  1.Wang ZW, Wen P, Qu YF, Dong SH, Li JJ, Tan K, Nieh JC: Bees eavesdrop upon informative and persistent signal compounds in alarm pheromones. Scientific Reports 2016, 6. 

  2.Wang ZW, Qu YF, Dong SH, Wen P, Li JJ, Tan K, Menzel R: Honey Bees Modulate Their Olfactory Learning in the Presence of Hornet Predators and Alarm Component. PLoS One 2016, 11(2). 

  3.Tan K, Qu YF, Wang ZW, Liu ZW, Engel MS: Haplotype diversity and genetic similarity among populations of the Eastern honey bee from Himalaya-Southwest China and Nepal (Hymenoptera: Apidae). Apidologie 2016, 47(2):197-205. 

  4.Tan K, Dong SH, Li XY, Liu XW, Wang C, Li JJ, Nieh JC: Honey Bee Inhibitory Signaling Is Tuned to Threat Severity and Can Act as a Colony Alarm Signal. Plos Biology 2016, 14(3). 

  5.Gong ZW, Wang C, Nieh JC, Tan K: Inhibiting DNA methylation alters olfactory extinction but not acquisition learning in Apis cerana and Apis mellifera. J Insect Physiol 2016, 90:43-48. 

  6.Wen P, Mo JC, Lu CW, Tan K, Sobotnik J, Sillam-Dusses D: Sex-pairing pheromone of Ancistrotermes dimorphus (Isoptera: Macrotermitinae). J Insect Physiol 2015, 83:8-14. 

  7.Tan K, Wang YC, Dong SH, Liu XW, Zhuang D, Chen WW, Oldroyd BP: Associations between reproduction and work in workers of the Asian hive bee Apis cerana. J Insect Physiol 2015, 82:33-37. 

  8.Tan K, Liu XW, Dong SH, Wang C, Oldroyd BP: Pheromones affecting ovary activation and ovariole loss in the Asian honey bee Apis cerana. J Insect Physiol 2015, 74:25-29. 

  9.Tan K, Latty T, Dong SH, Liu XW, Wang C, Oldroyd BP: Individual honey bee (Apis cerana) foragers adjust their fuel load to match variability in forage reward. Scientific Reports 2015, 5. 

  10.Tan K, Dong SH, Liu XW, Chen WW, Wang YC, Oldroyd BP, Latty T: Phantom alternatives influence food preferences in the eastern honeybee Apis cerana. J Anim Ecol 2015, 84(2):509-517. 

  11.Tan K, Chen WW, Dong SH, Liu XW, Wang YC, Nieh JC: A neonicotinoid impairs olfactory learning in Asian honey bees (Apis cerana) exposed as larvae or as adults. Scientific Reports 2015, 5. 

  12.Holmes MJ, Tan K, Wang Z, Oldroyd BP, Beekman M: Genetic reincarnation of workers as queens in the Eastern honeybee Apis cerana. Heredity 2015, 114(1):65-68. 

  13.Wang ZW, Tan K: Comparative analysis of olfactory learning of Apis cerana and Apis mellifera. Apidologie 2014, 45(1):45-52. 

  14.Wang ZW, Chen G, Tan K: Both olfactory and visual cues promote the hornet Vespa velutina to locate its honeybee prey Apis cerana. Insectes Sociaux 2014, 61(1):67-70. 

  15.Tan K, Latty T, Hu ZWN, Wang ZW, Yang S, Chen WW, Oldroyd BP: Preferences and tradeoffs in nectar temperature and nectar concentration in the Asian hive bee Apis cerana. Behavioral Ecology and Sociobiology 2014, 68(1):13-20. 

  16.Tan K, Chen WW, Dong SH, Liu XW, Wang YC, Nieh JC: Imidacloprid Alters Foraging and Decreases Bee Avoidance of Predators. PLoS One 2014, 9(7). 

  17.Remnant EJ, Koetz A, Tan K, Hinson E, Beekman M, Oldroyd BP: Reproductive interference between honeybee species in artificial sympatry. Mol Ecol 2014, 23(5):1096-1107. 

  18.Li JJ, Wang ZW, Tan K, Qu YF, Nieh JC: Effects of natural and synthetic alarm pheromone and individual pheromone components on foraging behavior of the giant Asian honey bee, Apis dorsata. Journal of Experimental Biology 2014, 217(19):3512-3518. 

  19.Li JJ, Wang ZW, Tan K, Qu YF, Nieh JC: Giant Asian honeybees use olfactory eavesdropping to detect and avoid ant predators. Animal Behaviour 2014, 97:69-76. 

  20.Holmes MJ, Tan K, Wang Z, Oldroyd BP, Beekman M: Why acquiesce? Worker reproductive parasitism in the Eastern honeybee (Apis cerana). Journal of Evolutionary Biology 2014, 27(5):939-949. 

  21.Holmes MJ, Tan K, Wang Z, Oldroyd BP, Beekman M: Effect of queen excluders on ovary activation in workers of the Eastern honeybee Apis cerana. Insectes Sociaux 2014, 61(2):191-196.

22.Tan K, Yang S, Wang Z, Menzel R: Effect of Flumethrin on Survival and Olfactory Learning in Honeybees. PLoS One 2013, 8(6). 

23. Tan K, Wang Z, Chen W, Hu Z, Oldroyd BP: The 'I see you' prey-predator signal of Apis cerana is innate. Naturwissenschaften 2013, 100(3):245-248. 

24.  Tan K, Hu Z, Chen W, Wang Z, Wang Y, Nieh JC: Fearful Foragers: Honey Bees Tune Colony and Individual Foraging to Multi-Predator Presence and Food Quality. PLoS One 2013, 8(9). 

25.  Holmes MJ, Tan K, Wang Z, Oldroyd BP, Beekman M: Honeybee (Apis cerana) guards do not discriminate between robbers and reproductive parasites. Insect Soc 2013, 60(2):265-271. 

26. Tan K, Yang M-X, Wang Z-W, Li H, Zhang Z-Y, Radloff SE, Hepburn R: Cooperative wasp-killing by mixed-species colonies of honeybees, Apis cerana and Apis mellifera. Apidologie 2012, 43(2):195-200. 

27.  Tan K, Yang M, Wang Z, Radloff SE, Pirk CWW: The pheromones of laying workers in two honeybee sister species: Apis cerana and Apis mellifera. J Comp Physiol A -Neuroethol Sens Neural Behav Physiol 2012, 198(4):319-323. 

28. Tan K, Wang Z, Yang M, Fuchs S, Luo L, Zhang Z, Li H, Zhuang D, Yang S, Tautz J et al: Asian hive bees, Apis cerana, modulate dance communication in response to nectar toxicity and demand. Anim Behav 2012, 84(6):1589-1594. 

 29.  Tan K, Wang Z, Li H, Yang S, Hu Z, Kastberger G, Oldroyd BP: An 'I see you' prey-predator signal between the Asian honeybee, Apis cerana, and the hornet, Vespa velutina. Anim Behav 2012, 83(4):879-882. 

 30.T.Ken, Fuchs S. and Ruiguang Z.2003, Morphometrical characterization of Apis cerana in   Yunnan  Province of China. Apidologie,34(6)553-562.

31.  T.Ken, H.R. Hepburn, S.E. Radloff, 2005,Heat-balling to Vespa velutina , Naturwissenshaften,92:491-495.

32.   T.Ken,2006,Gigantism:rearing Apis cerana queen in mixed  colonies, Naturwissenschaften 93:315-320

33.  T.Ken,2, Marina D. Meixner, Stefan Fuchs , 2006, Morphological and Molecular Studies     of Apis cerana Fabr. (Apidae, Hymenoptera) in China. Systematics and Biodiversity 4 (4): 473–482

34.  T.Ken, Natapot W,Deborah Roan S,2007, Mitochondrial DNA diversity of Chinese Apis cerana,Apidologie,36(3)238-246. 

35. T.Ken,2007, Environmentally-induced developmental effects on morphometric characters of workers in Apis cerana colonies .Apidologie,36(3)289-295.

36.  T.Ken,2007,Bee-hawking by the wasp, Vespa velutina, on the honeybeesApis cerana and A. Mellifera, Naturwissenschaften 94 469-472

37. T.Ken , Y. H. Guo,S. W. Nicolson, S. E. Radloff ,Q. S. Song , H. R. Hepburn Honeybee (Apis cerana) Foraging Responses to the Toxic Honey of Tripterygium hypoglaucum (Celastraceae):Changing Threshold of Nectar Acceptability, Journal of Chemical Ecology (2007) 33:2209–2217

38.   T.Ken,  2008, Dancing to different tunes: heterospecific deciphering of the honeybee waggle dance    Naturwissenschaften, 95:1165-1168

39.  T.Ken, H.R. Hepburn,S.E. Radloff., 2008, Mutiple analysis of China Apis cerana,Apidologie, 39(3)343-353

40.  T.Ken,Stefan Fuchs,Michael Engel,2008, An adventitious distal abscissa in the forewing of honey bees, Apidologie, 39(6)674-682

41.  T. Ken , H.R. Hepburn, 2009, Intra-and interspecific brood Recognition in honeybee Apis cerana and Apis mellifera, Apidologie,40(2)184-191

42.  T. Ken, M.X.Yang, H.R. Hepburn,S.E. Radloff.(2009) Worker reproduction in mixed-species colonies of honey bees,Behavioural Ecology,20(5):1106-1110

43.  B. Dainat, T. Ken,*, H. Berthoud, P. Neumann,2009, The ectoparasitic mite Tropilaelaps mercedesae (Acari, Laelapidae) as a vector of honeybee viruses,  Insect Sociaux,56 (2009) 40 – 43

44.  Yang MX, Tan K*, Radloff SE, Pirk CWW, Hepburn HR,2010, Hetero-specific queen retinue behavior of worker bees in mixed-species colonies of Apis cerana and Apis mellifera. Apidologie 41:54-61

45.  T. Ken, H. Li, M.X. Yang, H.R. Hepburn, S.E. Radloff,, 2010, Wasp hawking induces endothermic heat production in guard bees, Journal of Insect Science, 10(142): 1-6

46.  T. Ken,Z.W.Wang, Hua Li, M.X. Yang. C. W. W. Pirk, H. Randall,Sarah E. Radloff, 2010, Responses of Queenright and Queenless Workers of Apis Cerana to 9-keto-2(E)-decenoic Acid, a Pheromonal Constituent of the Mandibular Gland, Journal of Chemical ecology,36:966-968

47.   T. Ken, Z.W.Wang, M.X. Yang, H.R. Hepburn, S.E. Radloff, 2010, Nestmate Recognition Differences between Honeybee Colonies of Apis cerana and Apis mellifera, Journal of insect behavior, 2010, 23:311-388

48.  T. Ken, M.X. YANG., Z.W. WANG.. H. LI.,, Z.Y. ZHANG.,, Sarah E. RADLOFF.* and Randall HEPBURN. Cooperative wasp-killing by mixed-species colonies of honeybees, Apis cerana and Apis mellifera Apidologie,43(2) 195-200

49.  T. Ken, Z.W.Wang, M.X. Yang, H.R. Hepburn, S.E. Radloff, 2012, The pheromones of  workers in two honeybees sister species: Apis cerana and Apis mellifera, Journal of comparative physiology, 198:319-323

50.  T. Ken, Z.W.Wang, H. Li, G.Kasburg, B, Oldroyd, 2012,An “I see you” prey to predator signal between the Asian honeybee (Apis cerana) and the hornet (Vespa velutina) Journal of Animal Behaviour, 83(2012)879-882

51.T. Ken,, M.X. YANG., Z.W. WANG.. H. LI.,, Z.Y. ZHANG.,, Sarah E. RADLOFF.* and Randall HEPBURN,Commence foraging temperature between Apis cerana and Apis mellifera, Apidologie,43(6) 618-623

52.T.Ken, Wang.Z.W,et.al 2012, Asian hive bees, Apis cerana, modulate dance communication in response to nectar toxicity and demand, Journal of Animal Behaviour, 83(2012) 1589-1594

53. Ting ZHOU, Denis L ANDERSON, Zachary Y HUANG , Ken TAN, 2004, Identification of Varroa mites infesting Apis cerana and Apis mellifera in      China.Apidologie,34(6)553-562,

54. S.E. RADLOFF, H.R. HEPBURN, Ken TAN ,2004,. Multivariate morphometric analysis of of southern mainland Asia Apidologie,35(1)475-492

55. H.R. Hepburn, S.E. Radloff, G.W. Otis, S. Fuchs, Ken Tan,2005, Multivariate morphometrica nalysis of Apis florea,Apidologie, 36(3)359-376

56. S.E. RADLOFF, H.R. HEPBURN, S. FUCHS, Ken Tan,2005,. Multivariate morphometric analysis of Apis cerana populations of oceanic Asia.. Apidologie, 36(3)127-140

57. Jianjun Li, Ken Tan,and Shaoyu He,2008, Hornet predation risk: apperception and response by honey bees,Journal of apicultural research, 47(1): 174–175

58.M.X. Yang, S.E. Radloff, K. Tan, H.R. Hepburn 2010, Anti-predator Fan-blowing in Guard Bees, Apis mellifera capensis Esch,,Journal of insect behaviour, 23:12–18

59. M.X.Yang, Z.W. Wang, H Li, Zu-Yun Zhang, Ken Tan 2010, Thermoregulation in mixed species colonies of honeybees (Apis cerana and Apis mellifera) ,Journal of insect physiology, 56, 706–709

60.  M.X.Yang ,K. Tan, S. E. Radloff , M. Phiancharoen, H. R. Hepburn,2010, Comb construction in mixed species colonies of honeybees Apis cerana and Apis mellifera,Journal of experimental of biology, 213, 1659-1664

61.  S. E. Radloff, C. Hepburn, H. R. Hepburn, S. Fuchs, S. Hadisoesilo, Ken Tan, M.S. Engel, V. Kuznetsov,2010, Population structure and classification of Apis cerana, Apidologie, 41:589-601

Community service:
  2004-present Chair of Bee Biology section of Asian Apicultural Association

Commitment to research the situation:
2001-2006    Genetic diversity of Yunnan honeybee, RMB100,000;
2006-2008   Breeding and selection of A.cerana, RMB80,000;
2006-2009  Breeding system of A. cerana, RMB 200,000;
2006-2009  Genetic diversity of China A. cerana, RMB260,000;

2013-2016  Genetic diversity of China feral honeybee  in Himalaya mountain region, RMB 470,000.