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TONG Haowen
   142 Xizhimenwai ST., Beijing, 100044, China
Zip Code  

Education and Appointments:

1978-1982: Got Bachelor Degree in geology in the Department of Geology in Northwest University in Xi’an, Shaanxi.

1982-1985: Got Master Degree in paleontology and stratigraphy in the Department of Geology in Northwest University in Xi’an, Shaanxi.

1985-1989: Got the Doctoral Degree in paleontology in the Department of Marine Geology in Tongji University in Shanghai.

1998: Visiting scholar at the Centre de Paléontologie stratigraphique et Paléoécologie, Université Claude Bernard - Lyon 1.

1999: Institute de Paléontologie Humaine, MNHN (Paris), postdoctoral fellowship

Research Professor in the Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences.

Adjunct Professor of the Graduate University of Chinese Academy of Sciences

Research Interest:
Quaternary mammalian fossils from prehistoric sites: their evolution and paleoenvironmental implications, mainly focusing on pachyderms; relationships between humans and the mammalian faunas.
Public Services:


Seleted Publication:

TONG Haowen, 1993: Eocene lacustrine deposits of China. Kaupia, Darmstadter Beitrage zur Naturgeschichte, Heft 2, 13-20.(in English)

TONG Haowen, 1995: Continuing diversifying--the only megatrend of evolution, Chinese Biodiversity , Vol.3, suppl., 94-96.(in English)

TONG Haowen, 1995: Overview of species concepts and remarks on intraspecific classification. Acta Palaeotologica Sinica, 34(6), 761--776. (in Chinese with English abstract)

TONG Haowen, 1997. Overview with some comments on speciation theories. Acta Palaeontologica Sinica, 36(3):387-400. (in Chinese with English summary)

TONG Haowen. 2000. Les Rhinocéros de sites à fossiles humains de Chine. L'Anthropologie. 104(2000) :523~529. (SSCI)

TONG Haowen, 2001a. Age profiles of rhino fauna from the Middle Pleistocene Nanjing Man Site, south China----Explained by the rhino specimens of living species. International Journal of Osteoarchaeology, 11(3): 231~237. (SSCI)

TONG Haowen, 2001b. Rhinocerotids in China----Systematics and material analysis. Geobios, 34(5):585-591.  (SCI)

TONG H.W., 2002. Java fauna: compared with that of Zhoukoudian area and South China. Acta Anthropologica Sinica 21, 325–336 (in Chinese with English abstract).

TONG H.W., 2003. Dicerorhinus mercki (Jager). In: Wu, R.K., et al. (Eds.), Homo erectus from Nanjing. Jiangsu Science and Technology Publishing House, Nanjing, pp. 111–120 (in Chinese with English summary).

TONG H.W., 2004a. Paleoenvironmental significance of Coelodonta in different fossil assemblages. Acta Anthropologica Sinica, 23 (suppl.), 306–314 (in Chinese with English abstract).

TONG H W, 2004b. Tapiridae, Chalicotheriidae. In: ZHENG S H (ed.), Jianshi Hominid Site. Beijing: Science Press, 233-253. (in Chinese with English summary)

TONG H.W., 2005a. Hystrix subcristata (Mammalia, Rodentia) from Tianyuan Cave, a human fossil site newly discovered near Zhoukoudian. Vertebrata PalAsiatica 43, 135–150 (in Chinese with English summary).

TONG Haowen. 2005b, Dental characters of the Quaternary tapirs in China, their significance in classification and phylogenetic assessment. Geobios, 38 : 139-150 (SCI)

TONG H.W, 2006a. Composition des faunes de mammifères quaternaires en Chine selon un gradient Nord-Sud. Lanthropologie,110 (2006) 870–887. (SSCI)

TONG H.W., 2006b. Hesperotherium sinense, a chalicothere (Perissodactyla, Mammalia) from the Early Pleistocene Liucheng Gigantopithecus Cave. Vertebrata PalAsiatica, 44(4), 347–365. (in Chinese with English summary)

TONG H W, 2007a. Aeretes melanopterus (Pteromyinae, Rodentia) from Tianyuan Cave near Zhoukoudian (Choukoutien) in China. Geobios, 40 (2007) 219–230. (SCI)

TONG Haowen, 2007b. Occurrences of warm-adapted mammals in north China over the Quaternary Period and their paleo-environmental significance. Science in China Series D: Earth Sciences, 2007, 50(9): 1327-1340. (SCI)

TONG H.W., 2008. Quaternary Hystrix (Rodentia, Mammalia) from North China: Taxonomy, stratigraphy and zoogeography, with discussions on the distribution of Hystrix in Palearctic Eurasia. Quaternary International, 179 (2008):126–134  (SCI)

Tong H W, Claude Guérin, 2009. Early Pleistocene Dicerorhinus sumatrensis remains from the Liucheng Gigantopithecus Cave, Guangxi, China. Geobios 42 (2009): 525–539.  (SCI)

TONG H W, LI H, XIE J Y, 2008. Revision of some taxa of the Salawusu Fauna from Sjara-osso-gol area, Nei Mengol, China. Quaternary Science, 28(6):1106-1113. (in Chinese with English summary)

TONG Haowen, LIU Jinyi, 2004. The Pleistocene-Holocene extinctions of mammals in China. Proceedings of the Ninth Annual Symposium of the Chinese Society of Vertebrate Paleontology , DONG Wei (ed). Beijing: China Ocean Press, 111~119. (in Chinese with English summary)

TONG Haowen, LIU Jinyi, HAN Ligang, 2002. On fossil remains of Early Pleistocene tapir (Perissodactyla, Mammalia) from Fanchang, Anhui. Chinese Science Bulletin, 47(7):586-590. (SCI)

TONG H W, LIU J Y, ZHANG S Q. 2004. Large mammal fossils from the Tianyuan Cave,a human fossil site of end pleistocene age recently discovered near Zhoukoudian. Acta Anthrop Sin (in Chinese with English summary), 2004, 23(3): 213―223

TONG H W, MOIGNE Anne-Marie, 2000. Quaternary rhinoceros of China. Acta Anthropologica Sinica, Supplement to vol. 19 (2000) : 257~263.

TONG H W, PATOU-MATHIS M.. 2003. Mammoth  and  other  proboscideans in China during the late Pleistocene. In Reumer J.W.F., de Vos J., Mol D. (eds), Advances in mammoth research. EINSEA,9 :421-428.

TONG H W, QIU Z X, 1991: An introduction to taphonomy.    Vertebrata PalAsiatica, 29(2),152-161. (in Chinese)

TONG H W, SHANG Hong, ZHANG Shuangquan & CHEN Fuyou,2004.A preliminary report on the newly found Tianyuan Cave, a Late Pleistocene human fossil site near Zhoukoudian. Chinese Science Bulletin, vol. 49, No.8: 853-857. (SCI)

TONG H W, SHANG H, ZHANG S Q, 2006. Mammalian Biostratigraphy of Tianyuan Cave, compared with that of Upper Cave at Zhoukoudian (Choukoutien). Acta Anthrop Sin, 2006, 25(1): 68―81. (in Chinese with English summary)

TONG H.W., XU F., 2001. On the origin and evolution of Quaternary tapirs in China. In: Wang,Y., Deng, T. (Eds.), Proceedings of the Eighth Annual Meeting of the Chinese Society ofVertebrate Paleontology. China Ocean Press, Beijing, pp. 133–141 (in Chinese with English abstract).

TONG H W, ZHANG S Q, CHEN F Y, LI Q, 2008. Rongements sélectifs des os par les porcs-épics et autres rongeurs : cas de la grotte Tianyuan, un site avec des restes humains fossiles récemment découvert près de Zhoukoudian (Choukoutien). L’anthropologie 112 (2008): 353–369  (SSCI)

TONG H W, ZHANG Shuang-quan, LI Qing, XU Zhi-jun, 2008. Late Pleistocene mammalian fossils from the Xitaiping Cave, Shidu, Beijing. Vertebrata PalAsiatica, 46(1), 51-70.  (in Chinese with English summary)

TONG H W, ZHANG Yonglu, 1994: How to use Latin names of taxa and their English equivalents. Vertebrata PalAsiatica, 32(2), 143-148.  (in Chinese with English summary)

JIN ChangZhu, QIN DaGong, PAN WenShi, TANG ZhiLu, LIU JinYi, WANG Yuan, DENG ChengLong, ZHANG YingQi, DONG Wei & TONG HaoWen, 2009. A newly discovered Gigantopithecus fauna from Sanhe Cave, Chongzuo, Guangxi, South China. Chinese Science Bulletin, 54 ( 5 ) : 788-797. (SCI)

LI Q, TONG H W, 2008. Age Structure Analysis of Cervus nippon in Tianyuan Cave at Zhoukoudian. Acta Anthropologica Sinica, 27(2):143-152.

LOUYS J, CURNOE D, TONG H., 2007. Characteristics of Pleistocene megafauna extinctions in Southeast Asia. Palaeogeography,  Palaeoclimatology, Palaeoecology, 243 (2007) 152–173 (SCI)

MADE van der J., TONG H.W.,2008. Phylogeny of the giant deer with palmate brow tines Megaloceros from west and Sinomegaceros from east Eurasia. Quaternary International, 179 (2008): 135–162  (SCI)

Supported Projects:

1995: Director’s Grants of IVPP (Grant number: 950801) : “Study on the new theory on bio-evolution”.

1997-1998: Special grants of the Chinese Academy of Sciences for the study of paleontology (Grant number: 970503) “Study of the Quaternary rhinocerotids from China”.

2000-2001: Special grants of China`s Ministry of Education for Returned Overseas Scholars (Grant number: KF200405): “The evolution of the diversity of Pleistocene rhinocerotids in China”.

2004-2006: NNSFC (National Natural Science Foundation of China) grants (Grant number: 40372015): “Research about the newly found Late Pleistocene fossil locality near Zhoukoudian  (Chou-kou-tien)”.

2005-2007: President’s Grants of the Chinese Academy of Sciences (Grant number: KL203302). “Excavation and research on the Tianyuan Cave, a newly discovered Late Pleistocene human site near Zhoukoudian (Chou-kou-tien)”.

2006-2011: Evolution of marine and terrestrial geobiodiversity during the earth history--Major Basic Research Projects (2006CB806400) of MST of China (Participate)

2007-2012: Investigation of the Chinese Anthropological Sites and resource, and the Collecting, Integrating to their Basic Data (2007FY110200) (Participate)